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Gryposuchus

Gryposuchus
Temporal range: Middle - Late Miocene, 13.65–7.25 Ma
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Crocodilia
Family: Gavialidae
Subfamily: Gryposuchinae
Genus: Gryposuchus
Gurich, 1912
Species
  • G. neogaeus (Burmeister, 1885 [originally Ramphostoma neogaeus]) (type)
  • G. jessei Gurich, 1912
  • G. colombianus Langston, 1965
  • G. croizati Riff and Aguilera, 2008
  • G. pachakamue Salas-Gismondi et al., 2016

Gryposuchus is an extinct genus of gavialoid crocodilian. It is the type genus of the subfamily Gryposuchinae. Fossils have been found from Argentina, Colombia, Venezuela, Brazil and the Peruvian Amazon. The genus existed during the early and middle Miocene epoch. One recently described species, G. croizati, grew to an estimated length of 10 metres (33 ft).

The type species of Gryposuchus is G. neogaeus. Specimens from this species were first described from Argentina in 1885, although it was referred to Ramphostoma at the time. It was assigned to the current genus in 1912 along with a newly described species, G. jessei.

Another species, G. colombianus, has been recovered from deposits in Colombia that date back to the Miocene and Pliocene epochs. This species, named in 1965, was originally referred to Gavialis. Fragmentary material of Gryposuchus from the Fitzcarrald Arch in the Peruvian Amazon dating back to the late middle Miocene bear a close resemblance to G. colombianus, but differ in rostrum proportions.

Some skull material also recovered from Peruvian Amazon (Iquitos) in the Pebas Formation, has been named as Gryposuchus pachakamue in 2016 by Roberto Salas-Gismondi et al. It includes the holotype MUSM 987, a well preserved skull that lacks of temporal and occipital bones; it measures 623.2 millimeters in length, and a series of referred specimens, including possible juveniles. The species was named after the Quechuan word for a primordial god and "storyteller". This new species is characterized by have 22 teeth in the mandible and the maxilla, a snout comparable in relative length to the modern Gavialis gangeticus, and is notable since that its orbits were wider than long and not so upturned as another species of gavialids, including the gryposuchines, which implies that G. pachakamue doesn't had the "telescoped" orbits (protruding eyes) condition fully developed. Since that it species, that inhabited the proto-Amazon fluvial system 13 million years ago, is the oldest record of gavialids in this area and it had a primitive telescoped eyes condition, it shows that the development of such condition was a case of convergent evolution with the species of Gavialis also found in fluvial environments.


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