Locusts are certain species of short-horned grasshoppers in the family Acrididae that have a swarming phase. These insects are usually solitary, but under certain circumstances, become more abundant and change their behaviour and habits, becoming gregarious. No taxonomic distinction is made between locust and grasshopper species; the basis for the definition is whether a species forms swarms under intermittently suitable conditions.
In the solitary phase, these grasshoppers are innocuous, their numbers are low, and they do not pose a major economic threat to agriculture. However, under suitable conditions of drought followed by rapid vegetation growth, serotonin in their brains triggers a dramatic set of changes: they start to breed abundantly, becoming gregarious and nomadic (loosely described as migratory) when their populations become dense enough. They form bands of wingless nymphs which later become swarms of winged adults. Both the bands and the swarms move around and rapidly strip fields and cause damage to crops. The adults are powerful fliers; they can travel great distances, consuming most of the green vegetation wherever the swarm settles.
Locusts have formed plagues since prehistory. The Ancient Egyptians carved them on their tombs and the insects are mentioned in the Bible and the Quran. Swarms have devastated crops and been a contributory cause of famines and human migrations. More recently, changes in agricultural practices and better surveillance of locations where swarms tend to originate, have meant that control measures can be used at an early stage. The traditional means of control are based on the use of insecticides from the ground or the air, but other methods using biological control are proving effective.