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Karyorelictea

Karyorelictea
Loxodes rostrum from page 267 of "American journal of physiology" (1898).jpg
Illustration of Loxodes rostrum
Scientific classification
Domain: Eukaryota
(unranked): SAR
(unranked): Alveolata
Phylum: Ciliophora
Subphylum: Postciliodesmatophora
Class: Karyorelictea
Corliss, 1974 
Orders

Karyorelictea is a class of ciliates.

The majority of karyorelict taxa are unculturable, although clonal lines of Loxodes have been developed.

According to Lynn (2008), the Karyorelictea class is divided into three orders:

These three orders were defined morphologically, and have been confirmed with molecular phylogenetics.

All ciliates possess multiple dimorphic nuclei which have separated the functions of protein synthesis, performed by the macronuclei or somatic-nuclei, and heritable DNA replication, performed by the micronuclei or germ-line nuclei. Members of the class Karyorelictea are characterised by an inability to replicate the macronucleus, a process called amitosis, which must be re-synthesized from the micronucleus every cell division.

The term karyorelict derives from the ancient greek (káruon), meaning "hard-shelled seed, or nucleus", and Latin , meaning 'abandoned', because of the inability of these ciliates to replicate the somatic nucleus.

An alternative genetic code is used by the nuclear genome of some karyorelictid ciliates (e.g., Parduczia sp.). This code corresponds to translation table 27 and involves the reassignment of three codons:


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