Ctenoid

The skin of most fishes are covered with scales. Scales vary enormously in size, shape, structure, and extent, ranging from strong and rigid armour plates in fishes such as shrimpfishes and boxfishes, to microscopic or absent in fishes such as eels and anglerfishes. The morphology of a scale can be used to identify the species of fish it came from.

Cartilaginous fishes (sharks and rays) are covered with placoid scales. Most bony fishes are covered with the cycloid scales of salmon and carp, or the ctenoid scales of perch, or the ganoid scales of sturgeons and gars. Some species are covered instead by scutes, and others have no outer covering on the skin.

Fish scales are part of the fish's integumentary system, and are produced from the mesoderm layer of the dermis, which distinguishes them from reptile scales. The same genes involved in tooth and hair development in mammals are also involved in scale development. The placoid scales of cartilaginous fishes are also called dermal denticles and are structurally homologous with vertebrate teeth. It has been suggested that the scales of bony fishes are similar in structure to teeth, but they probably originate from different tissue. Most fish are also covered in a protective layer of mucus (slime).

Placoid scales are found in the cartilaginous fishes: sharks, rays, and chimaeras. They are also called dermal denticles. Placoid scales are structurally homologous with vertebrate teeth ("denticle" translates to "small tooth"), having a central pulp cavity supplied with blood vessels, surrounded by a conical layer of dentine, all of which sits on top of a rectangular basal plate that rests on the dermis. The outermost layer is composed of vitrodentine, a largely inorganic enamel-like substance. Placoid scales cannot grow in size, but rather more scales are added as the fish increases in size.

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