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Athyriaceae

Athyrioideae
Illustration Athyrium filix-femina0.jpg
Athyrium filix-femina
Scientific classification
Kingdom: Plantae
(unranked): Polypodiophyta
Subclass: Polypodiidae
Order: Polypodiales
Family: Aspleniaceae
Subfamily: Athyrioideae
B.K.Nayar
Type genus
Athyrium
Genera

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See text

Athyrioideae is a subfamily of terrestrial ferns, with a cosmopolitan distribution.

Previously treated as a separate family, Athyriaceae Alston, in 2014 Christenhusz and Chase submerged it as subfamily Athyrioideae within family Aspleniaceae.

Originally considered as a member of the Eupolypods II clade, the Athyriaceae was related to other families in the clade as in this cladogram:

Cystopteridaceae

Rhachidosoraceae

Diplaziopsidaceae

Aspleniaceae

Hemidictyaceae

Thelypteridaceae

Woodsiaceae

Onocleaceae

Blechnaceae

Athyriaceae


The Athyriaceae in the past included Cystopteris and Gymnocarpium, now part of Cystopteridaceae. Inversely, this family has by some been subsumed in the family Woodsiaceae, but a Woodsiaceae defined in this way may be paraphyletic if it omits the Onocleaceae and Blechnaceae (as of 2006, the evidence was not clear).

Athyrioideae is placed within the Aspleniaceae as follows:

Cystopteridoideae (Acystopteris, Cystoathyrium, Cystopteris, Gymnocarpium)


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