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Valve (chiton)


Valve is an anatomical term applied to the shell of those molluscs that have a shell. Although in theory any mollusc shell can be termed a "valve", the word is now most commonly applied to members of two classes of molluscs: the Bivalvia (clams) and the Polyplacophora (chitons), in other words, to those molluscs whose shells are normally composed of more than one articulating part, each one of which is known as a valve (and also, in the case of chitons, as a "plate".)

Species within one family of very unusual small sea snails, marine opisthobranch gastropods in the family Juliidae, also have two articulating shells or valves, which resemble those of a bivalve. This exceptional family is commonly known as the bivalved gastropods. Gastropods in general are sometimes called "univalves", because in those that have a shell, the shell is usually in one part.

The valves of chitons are eight dorsal, shell plates, which are frequently coloured and sculpted. After death the girdle that holds the plates together disintegrates and the plates separate. Thus individual plates can be found washed up in beach drift, as shown in the image at the top of this article.

Bivalve molluscs (e.g. clams and oysters) have a shell which is composed of two separate but articulating parts. Each one of these two parts is known as a "valve".

The two valves are known as the "right valve" and the "left valve"; these are labeled with respect to the anterior end of the bivalve, when the umbones are facing upward. In many, but not all, bivalves, the two valves are more or less symmetrical and thus (other than the hinge line) look like mirror images of one another.

The great majority of shelled gastropods or snails have a shell in one part, hence the older name "univalve". The gastropod operculum, when present, even when it is composed mostly of calcium carbonate, is not considered to be a valve.


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