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Pholcidae

Cellar spiders
Pholcus.phalangioides.6905.jpg
Pholcus phalangioides
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Chelicerata
Class: Arachnida
Order: Araneae
Infraorder: Araneomorphae
Family: Pholcidae
C. L. Koch, 1851
Genera

See text.

Diversity
79 genera, 1461 species
Distribution.pholcidae.1.png
Estimated range of Pholcidae.

See text.

Pholcidae, commonly known as cellar spiders, are a spider family in the suborder Araneomorphae. The family contains about 1500 species divided into about 80 genera.

Some species, especially Pholcus phalangioides, are commonly called daddy long-legs spider, granddaddy long-legs spider, carpenter spider, daddy long-legger, or vibrating spider. Confusion often arises because the name "daddy long-legs" is also applied to two distantly related arthropod groups: the harvestmen (which are arachnids but not spiders), and crane flies (which are insects).

Pholcids are fragile arachnids, the body being 2–10 mm in length with legs which may be up to 50 mm long. Pholcus and Smeringopus have cylindrical abdomens and the eyes are arranged in two lateral groups of three and two smaller eyes. Eight and six eyes both occur in this family. Spermophora has a small abdomen and its eyes are arranged in two groups of three and no median eyes. Pholcids are gray to brown with banding or chevron markings. The shape of the Pholcus and Smeringopus's body resembles that of a peanut.

Pholcids are found in every continent in the world except Antarctica where it is too cold for them to survive. They hang inverted in messy, irregular, tangled webs. These webs are constructed in dark and damp recesses, such as: caves, under rocks and loose bark, abandoned mammal burrows, and undisturbed areas in buildings, such as cellars (hence the common name "cellar spider"). However, Pholcids are also quite commonly found in warm, dry places, such as household windows and attics.

The web has no adhesive properties but the irregular structure traps insects, making escape difficult. The spider quickly envelops its prey with silk and then inflicts the fatal bite. The prey may be eaten immediately or stored for later.

When the arachnid is threatened by a touch to the web or when too large a prey becomes entangled, the arachnid vibrates rapidly in a gyrating motion in its web and becomes blurred and difficult to focus on. For this reason pholcids have sometimes been called "vibrating spiders", although they are not the only species to exhibit this behavior. Doing so might make it difficult for a predator to see exactly where the spider is, may be intended to signal an assumed rival to leave, or may increase the chances of capturing insects that have just brushed their web and are still hovering nearby. If the spider continues to be harassed it will retreat into a corner or drop from its web and escape.


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