Maniraptor
| Maniraptorans Temporal range: Late Jurassic–Present, 167–0 Ma Possible Early Jurassic record |
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| Skeleton cast of Falcarius utahensis | |
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Clade: | Dinosauria |
| Order: | Saurischia |
| Suborder: | Theropoda |
| Clade: | Maniraptoriformes |
| Clade: |
Maniraptora Gauthier, 1986 |
| Subgroups | |
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| Synonyms | |
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Metornithes Perle et al., 1993 |
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Metornithes Perle et al., 1993
Maniraptora is a clade of coelurosaurian dinosaurs that includes the birds and the non-avian dinosaurs that were more closely related to them than to Ornithomimus velox. It contains the major subgroups Avialae, Deinonychosauria, Oviraptorosauria and Therizinosauria. Ornitholestes and the Alvarezsauroidea are also often included. Together with the next closest sister group, the Ornithomimosauria, Maniraptora comprises the more inclusive clade Maniraptoriformes. Maniraptorans first appear in the fossil record during the Jurassic Period (see Eshanosaurus), and are regarded as surviving today as living birds.
Maniraptorans are characterized by long arms and three-fingered hands (though reduced or fused in some lineages), as well as a "half-moon shaped" (semi-lunate) bone in the wrist (carpus). Maniraptorans are the only dinosaurs known to have breast bones (ossified sternal plates). In 2004, Tom Holtz and Halszka Osmólska pointed out six other maniraptoran characters relating to specific details of the skeleton. Unlike most other saurischian dinosaurs, which have pubic bones that point forward, several groups of maniraptorans have an ornithischian-like backwards-pointing hip bone. A backward-pointing hip characterizes the therizinosaurs, dromaeosaurids, avialans, and some primitive troodontids. The fact that the backward-pointing hip is present in so many diverse maniraptoran groups has led most scientists to conclude that the "primitive" forward-pointing hip seen in advanced troodontids and oviraptorosaurs is an evolutionary reversal, and that these groups evolved from ancestors with backward-pointing hips.
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