Makarkinia Temporal range: Aptian |
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Scientific classification | |
Kingdom: | Animalia |
Class: | Insecta |
Order: | Neuroptera |
Family: | †Kalligrammatidae |
Genus: |
†Makarkinia Martins-Neto, 1997 |
Species | |
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Makarkinia is an extinct genus of lacewing in the family Kalligrammatidae described by Martins-Neto in 1997 from a fossil found in South America. The genus contains two species dating from the Cretaceous, Makarkinia adamsi and Makarkinia kerneri.
When first described, Makarkinia was known from a single fossil wing which is a compression-impression fossil preserved in layers of soft sedimentary rock. Along with other well-preserved insect fossils, the Makarkinia specimens were collected from layers of the Upper Aptian Crato Formation. The formation is composed of unweathered grey and oxidized yellow limestones, which preserved numerous insects, fish, birds and reptiles as a notable lagerstätten. The area is a preserved inland lake or one of a series of lakes, though the nature as a fresh or salt-water body is uncertain. The depth of the basin has been suggested as either shallow or fairly deep. The basin formed near the center of the supercontinent Gondwana during the early part of the diversification of flowering plants.
The M. adamsi holotype specimen was preserved in the National Museum of Brazil paleontological collection when first studied. The fossil was described by R. G. Martins-Neto in a 1992 paper on the neuropterans of the Crato Formation, with the species named in it. At the time of description the species was placed into the extinct family Panfiloviidae as Panfilovia adamsi. This placement was changed in a 1997 by Martins-Neto who moved the species to a new genus, Makarkinia and new subfamily "Makarkiniinae" in Panfiloviidae. Three years later Martins-Neto elevated the subfamily to a full family as Makarkiniidae, though this change was not widely used. It was suggested by Makarkin and Archibald in 2003 and subsequent authors that the genus was closely related to Kalligrammatidae, and moved into the family in 2016 by Günter Bechly and Vladimir Makarkin.