Ilex | |
---|---|
Ilex paraguariensis | |
Scientific classification | |
Kingdom: | Plantae |
(unranked): | Angiosperms |
(unranked): | Eudicots |
Order: | Aquifoliales |
Family: |
Aquifoliaceae DC. ex A.Rich. |
Genus: |
Ilex L. |
Species | |
About 600, see text |
About 600, see text
Ilex /ˈaɪlɛks/, or holly, is a genus of 400 to 600 species of flowering plants in the family Aquifoliaceae, and the only living genus in that family. The species are evergreen or deciduous trees, shrubs, and climbers from tropics to temperate zones worldwide.
The genus Ilex is widespread throughout the temperate and subtropical regions of the world. It includes species of trees, shrubs, and climbers, with evergreen or deciduous foliage and inconspicuous flowers. Its range was more extended in the Tertiary period and many species are adapted to laurel forest habitat. It occurs from sea level to more than 2,000 metres (6,600 ft) with high mountain species. It is a genus of small, evergreen trees with smooth, glabrous, or pubescent branchlets. The plants are generally slow-growing with some species growing to 25 m (82 ft) tall. The type species is the European holly Ilex aquifolium described by Linnaeus.
Plants in this genus have simple, alternate glossy leaves, frequently with a spiny leaf margin. The inconspicuous flower is greenish white, with four petals. They are generally dioecious, with male and female flowers on different plants.
The small fruits of Ilex, although often referred to as berries, are technically drupes. They range in color from red to brown to black, and rarely green or yellow. The "bones" contain up to ten seeds each. Some species produce fruits parthenogenetically, such as the cultivar 'Nellie R. Stevens'. The fruits ripen in winter and thus provide winter colour contrast between the bright red of the fruits and the glossy green evergreen leaves. Hence the cut branches, especially of I. aquifolium, are widely used in Christmas decoration. The fruits are generally slightly toxic to humans, and can cause vomiting and diarrhea when ingested. However, they are an important food source for birds and other animals, which help disperse the seeds. Unfortunately this can have negative impacts as well. Along the west coast of North America, from California to British Columbia, English Holly (Ilex aquifolium), which is grown commercially, is quickly spreading into native forest habitat, where it thrives in shade and crowds out native species. It has been placed on the Washington State Noxious Weed Control Board's monitor list, and is a Class C invasive plant in Portland.