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Hadramphus spinipennis

Hadramphus spinipennis
Spinipennis.jpg
Coxella weevil on Aciphylla dieffenbachii

Nationally Vulnerable (NZ TCS)
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Family: Curculionidae
Subfamily: Molytinae
Genus: Hadramphus
Species: H. spinipennis
Binomial name
Hadramphus spinipennis
Broun, 1911

Hadramphus spinipennis, commonly called the coxella weevil, is a large, nocturnal, flightless weevil only found on Mangere and Rangatira Islands in the Chatham Islands, New Zealand.

Hadramphus spinipennis species was described by Thomas Broun as having an brown egg-shaped body with glossy brownish-black antennae. It has an prominent tubercle or wart-like projection on each side of its thorax and further tubercles on its abdomen, and a black underside. The length of the adult beetle ranges from 21–22 mm (females can reach 25 mm) excluding the rostrum, and its width is 10 mm. It is the only large weevil on the Chatham Islands.

This species was discovered by Thomas Hall, a sheep musterer on Pitt Island in the Chathams, who collected numerous specimens of undescribed species for Broun. Despite being described from Pitt Island, it has not been seen there since Hall discovered it, probably because its host plant there has been almost wiped out by sheep. It is one of four Hadramphus weevil species, all endemic to New Zealand, specialised, flightless, and rare.

This weevil's host plant is Aciphylla dieffenbachii Kirk (coxella or Dieffenbach's speargrass), a soft-leaved relative of the spine-bearing alpine Aciphylla of the New Zealand mainland. Adults make a characteristic oval feeding notch on the leaf petiole, sometimes scraping away the leaf to encourage to production of gum. Green seeds and flowers of coxella are eaten, with a preference for male flowers. Leaves are also gummed together to shelter a feeding larva, but larve also feed in the crown of the plant amongst leaf bases, and if several are present they can kill the plant.

Because they sometimes kill their host plants, adult weevils are able to walk considerable distances – up to 600 m – in search of new patches of coxella. They seem to form a metapopulation, relying on the constant re-establishment of new patches of Aciphylla after old patches are wiped out by high weevil densities. Coxella weevils are also regularly found on Chatham Island lancewood (Pseudopanax chathamicum), on which they seem to shelter but only intermittently eat.


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Wikipedia

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