Guttiferae

Clusiaceae
Clusia major
Scientific classification
Kingdom:	Plantae
(unranked):	Angiosperms
(unranked):	Eudicots
(unranked):	Rosids
Order:	Malpighiales
Family:	Clusiaceae Lindl.
Type genus
Clusia

The Clusiaceae or Guttiferae Juss. (1789) (nom. alt. et cons. = alternative and valid name) are a family of plants including 13 genera and ca 750 species,. Several former members of Clusiacae are now placed in Calophyllaceae and Hypericaceae. They are mostly trees and shrubs, with milky sap and fruits or capsules for seeds. The family is primarily tropical. More so than many plant families, it shows large variation in plant morphology (for example, three to 10, fused or unfused petals, and many other traits). According to the APG III, this family belongs to the order Malpighiales.

One feature which is sometimes found in this family, and rarely in others (e.g., Malpighiaceae), is providing pollinators with rewards other than pollen or nectar; specifically, some species offer resin which bees use in nest construction (all three rewards are found in different species of the Clusiaceae).

The family Clusiaceae was divided by Cronquist into two subfamilies: the Clusioideae (typical subfamily) and the Hypericoideae. The latter was often treated as a family - the Hypericaceae or St. John's wort family. Elements of the Hypericoideae are more common in northern temperate areas and those of the Clusioideae are centered in the tropics.

Later classifications, however, divide the family in a finer way. Molecular studies have shown that the family Podostemaceae - the riverweeds - as well as the Bonnetiaceae are nested in this group. Their inclusions make the Clusiaceae in a wide-sense polyphyletic, and Stevens's subfamilies need to be recognised at family level: Clusioideae as Clusiaceae sensu stricto; Hypericoideae as Hypericaceae; and Kielmeyeroideae as Calophyllaceae.

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