Coleoptera

Beetle Temporal range: 318–0 Ma PreЄ Є O S D C P T J K Pg N Late Carboniferous–Holocene
Clockwise from top left: female golden stag beetle (Lamprima aurata), rhinoceros beetle (Megasoma sp.), long nose weevil (Rhinotia hemistictus), cowboy beetle (Chondropyga dorsalis), and a species of Amblytelus.
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
(unranked):	Endopterygota
Order:	Coleoptera Linnaeus, 1758
Suborders
Adephaga Archostemata Myxophaga Polyphaga † See subgroups of the order Coleoptera

See subgroups of the order Coleoptera

Beetles are a group of insects that form the order Coleoptera, in the superorder Endopterygota. Their front pair of wings is hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 species, is the largest of all orders, constituting almost 40% of described insects and 25% of all known animal life-forms; new species are discovered frequently. The largest of all families, the Curculionidae (weevils) with some 70,000 member species, belongs to this order. They are found in almost every habitat except the sea and the polar regions. They interact with their ecosystems in several ways: beetles often feed on plants and fungi, break down animal and plant debris, and eat other invertebrates. Some species are serious agricultural pests, such as the Colorado potato beetle, while others such as Coccinellidae (ladybirds or ladybugs) eat aphids, scale insects, thrips, and other plant-sucking insects that damage crops.

Beetles typically have a particularly hard exoskeleton including the elytra, though some such as the rove beetles have very short elytra while blister beetles have softer elytra. The general anatomy of a beetle is quite uniform and typical of insects, although there are several examples of novelty, such as adaptations in water beetles which trap air bubbles under the elytra for use while diving. Beetles are endopterygotes, which means that they undergo complete metamorphosis, with a series of conspicuous and relatively abrupt changes in body structure between hatching and becoming adult after a relatively immobile pupal stage. Some, such as, stag beetles have a marked sexual dimorphism, the males possessing enormously enlarged mandibles which they use to fight other males. Many beetles are aposematic, with bright colours and patterns warning of their toxicity, while others are harmless Batesian mimics of such insects. Many beetles, including those that live in sandy places, have effective camouflage.