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Chrysochlorinae

Golden moles
Temporal range: Lutetian–Recent
Taupe doree.jpg
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Afrosoricida
Suborder: Chrysochloridea
Broom, 1915
Family: Chrysochloridae
Gray, 1825
Genera

Golden moles are small, insectivorous burrowing mammals endemic to Southern Africa, where their Afrikaans names are gouemolle or kruipmolle (singular gouemol or kruipmol). They comprise the family Chrysochloridae and as such they are taxonomically distinct from the true moles, family Talpidae, and other mole-like families, all of which, to various degrees, they resemble as a result of convergence.

Like most burrowing mammals with similar habits, the Chrysochloridae have short legs with powerful digging claws, very dense fur that repels dirt and moisture, and toughened skin, particularly on the head. Their eyes are non-functional and covered with furred skin. The external ears are just tiny openings. In particular, golden moles bear a remarkable resemblance to the marsupial moles of Australia, family Notoryctidae, which they resemble so suggestively that at one time, the marsupial/placental divide notwithstanding, some workers argued that they were related. Considerations that influenced the debate might have included the view that the Chrysochloridae are very primitive placentals and the fact that they have many mole-like specializations similar to specializations in marsupial moles.The rhinarium is a greatly enlarged, dry leathery pad that protects their nostrils while the animal digs. In this respect too, they resemble the marsupial moles. Some authors claim that their primary sense is that of touch, and they are particularly sensitive to vibrations that may indicate approaching danger. Note below however, the observations on the malleus in the middle ear.

The species range in size from about 8 to about 20 cm. They have muscular shoulders and the forelimbs are radically adapted for digging; all the toes on the forefeet have been reduced, except for a large, pick-like third claw on the third toe. The fifth digit is absent and the first and fourth digits are vestigial. The adaptations of the hind feet are less dramatic, they retain all five toes and are webbed as an adaptation to efficient backward shovelling of soil loosened by the front claws.


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