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Chironomidae

Chironomidae
Chironomus plumosus01.jpg
Male Chironomus plumosus
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Diptera
Suborder:
Infraorder: Culicomorpha
Superfamily: Chironomoidea
Family: Chironomidae
Genera

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The Chironomidae (informally known as chironomids, nonbiting midges, or lake flies) comprise a family of flies with a global distribution. They are closely related to the Ceratopogonidae, Simuliidae, and Thaumaleidae. Many species superficially resemble mosquitoes, but they lack the wing scales and elongated mouthparts of the Culicidae.

The name Chironomidae stems from the Ancient Greek word kheironómos, "a pantomimist").

This is a large taxon of insects; some estimates of the species numbers suggest well over 10000 world-wide. Males are easily recognized by their antennae. Adults are known by a variety of vague and inconsistent common names, largely by confusion with other insects. For example, chironomids are known as "lake flies" in parts of Canada and Lake Winnebago, Wisconsin, but "bay flies" in the areas near the bay of Green Bay, Wisconsin. They are called "sand flies", "muckleheads", "muffleheads", "Canadian soldiers", or "American soldiers" in various regions of the Great Lakes area. They have been called "blind mosquitoes" or "chizzywinks" in Florida. However, they are not mosquitoes of any sort, and the term "sandflies" generally refers to various species of biting flies unrelated to the Chironomidae.

The group includes Belgica antarctica, the largest terrestrial animal of Antarctica.

The biodiversity of the Chironomidae often goes unnoticed because they are notoriously difficult to identify and ecologists usually record them by species groups. Each morphologically distinct group comprises a number of morphologically identical (sibling) species that can only be identified by rearing adult males or by cytogenetic analysis of the polytene chromosomes. Polytene chromosomes were originally observed in the larval salivary glands of Chironomus midges by Balbiani in 1881. They form through repeated rounds of DNA replication without cell division, resulting in characteristic light and dark banding patterns which can be used to identify inversions and deletions which allow species identification.


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