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Chara (alga)

Chara
Temporal range: Silurian–Recent
CharaFragilis.jpg
Chara globularis
Scientific classification
Kingdom: Plantae
Division: Charophyta
Class: Charophyceae
Order: Charales
Family: Characeae
Genus: Chara
L., 1753
Species

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Chara is a genus of charophyte green algae in the family Characeae. They are multicellular and superficially resemble land plants because of stem-like and leaf-like structures. They are found in fresh water, particularly in limestone areas throughout the northern temperate zone, where they grow submerged, attached to the muddy bottom. They prefer less oxygenated and hard water and are not found in waters where mosquito larvae are present. They are covered with calcium carbonate deposits and are commonly known as stoneworts. Cyanobacteria have been found growing as epiphytes on the surfaces of Chara, where they may be involved in fixing nitrogen, which is important to plant nutrition.

The branching system of Chara species is complex with branches derived from apical cells which cut off segments at the base to form nodal and internodal cells alternately. The main axes bear whorls of branches in a superficial resemblance to Equisetum. They are typically anchored to the littoral substrate by means of branching underground rhizoids. Chara plants are rough to the touch because of deposited calcium salts on the cell wall. The metabolic processes associated with this deposition often give Chara plants a distinctive and unpleasant smell of hydrogen sulfide.

The plant body is a gametophyte. It consists of a main axis (differentiated into nodes and internodes), dimorphic branches (long branch of unlimited growth and short branches of limited growth), rhizoids (multicellular with oblique septa) and stipulodes (needle shaped structures at the base of secondary laterals).

Chara reproduces vegetatively and sexually. Vegetative reproduction takes place by tubers, amylum stars and secondary protonema. The fructifications for sexual reproduction are globule or antheridium (male) and nucule or archegonium (female). The sex organs are multicellular and jacketed. The antheridia and archegonia may occur on separate plants (dioecy), together on the same plant (conjoined monoecy) or separately on the same plant (sejoined monoecy). After fertilization, the zygote develops into an oospore.


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