Beeswax (cera alba) is a natural wax produced by honey bees of the genus Apis. The wax is formed into "scales" by eight wax-producing glands in the abdominal segments of worker bees, who discard it in or at the hive. The hive workers collect and use it to form cells for honey-storage and larval and pupal protection within the beehive. Chemically, beeswax consists mainly of esters of fatty acids and various long-chain alcohols.
Beeswax has long-standing applications in human food and flavoring. For example, it is used as a glazing agent, a sweetener, or as a light/heat source. It is edible, in the sense of having similar negligible toxicity to plant waxes, and is approved for food use in most countries and the European Union under the E number E901. However, the wax monoesters in beeswax are poorly hydrolysed in the guts of humans and other mammals, so they have insignificant nutritional value. Some birds, such as honeyguides, can digest beeswax. Beeswax is the main diet of Wax moth larvae.
The wax is formed by worker bees, which secrete it from eight wax-producing mirror glands on the inner sides of the sternites (the ventral shield or plate of each segment of the body) on abdominal segments 4 to 7. The sizes of these wax glands depend on the age of the worker, and after many daily flights, these glands begin to gradually atrophy.
The new wax is initially glass-clear and colourless, becoming opaque after mastication and adulteration with pollen by the hive worker bees. Also, the wax becomes progressively more yellow or brown by incorporation of pollen oils and propolis. The wax scales are about 3 mm (0.12 in) across and 0.1 mm (0.0039 in) thick, and about 1100 are required to make a gram of wax.