Azhdarchids Temporal range: Late Cretaceous, 108–66 Ma Possibly earlier |
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Reconstructed skeleton of Quetzalcoatlus northropi | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | †Pterosauria |
Suborder: | †Pterodactyloidea |
Clade: | †Neoazhdarchia |
Family: |
†Azhdarchidae Nesov, 1984 |
Type species | |
†Azhdarcho lancicollis Nesov, 1984 |
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Genera | |
See text |
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Synonyms | |
"Titanopterygiidae" |
See text
"Titanopterygiidae"
Padian, 1984 (preoccupied)
Azhdarchidae (from Persian word azhdar (اژدر), a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the early Cretaceous as well (late Berriasian age, about 140 million years ago). Azhdarchids included some of the largest known flying animals of all time, but members no larger than a cat have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the azhdarchinae to include the pterosaurs Azhdarcho, Quetzalcoatlus, and "Titanopteryx" (now known as Arambourgiania). They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. By the time of the end-Cretaceous mass extinction, most pterosaur families except for the Azhdarchidae disappear from the fossil record, but recent studies indicate a wealth in pterosaurian faunas, including pteranodontids, nyctosaurids, tapejarids and several indeterminate forms. Some taxa like Navajodactylus, Bakonydraco and Montanazhdarcho were moved from Azhdarchidae to other clades.
Azhdarchids are characterized by their long legs and extremely long necks, made up of elongated neck vertebrae which are round in cross section. Most species of azhdarchids are still known mainly from their distinctive neck bones and not much else. The few azhdarchids that are known from reasonably good skeletons include Zhejiangopterus and Quetzalcoatlus. Azhdarchids are also distinguished by their relatively large heads and long, spear-like jaws. There are two major types of azhdarchid morphologies: the "blunt-beaked" forms with shorter and deeper bills and the "slender-beaked" forms with longer and thinner jaws.