Sir Arthur Acland (also recorded as Sir Arthure Akelane) (1573–1610) of Acland in the parish of Landkey, Devon, was a member of the Devonshire gentry, and was knighted in 1606. Little is known of his life and career, but his monumental inscription survives above his impressive monument in Landkey Church. His son was Sir John Acland, 1st Baronet (c. 1591 – 1647). He was ancestor to the prominent, wealthy and long-enduring Acland family of Killerton, which survives today in the direct male line.
Arthur Acland's grandfather was John Acland (died 1553), of Acland, described as "the first of the [Acland] family to emerge from the shadows of history as a visible human being". His father was Hugh Acland (c. 1543 – 1622) of Acland, Sheriff of Devon in 1611, and he was the eldest of his four sons. His mother was Margaret Monke (died 1619), who was a daughter of Thomas Monke (c. 1515 – c. 1583), of Potheridge, who came from an ancient Devonshire family and was great-grandfather of George Monck, 1st Duke of Albemarle (1608–1670). Thomas Monke's first wife and Margaret's mother was Frances Plantagenet, a daughter of Arthur Plantagenet, 1st Viscount Lisle (died 1542) an illegitimate son of King Edward IV.
Acland married his step-first cousin Eleanor Mallet (1573–1645), a daughter and co-heiress of Robert Mallet of Woolleigh, Beaford. Her mother (or step-mother) was Elizabeth Rolle, who re-married to Sir John Acland (died 1620), Sir Arthur's uncle. Eleanor survived her husband and remarried to Sir Francis Vincent, 1st Baronet (c. 1568–1640) of Stoke d'Abernon, in Surrey.