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Typothorax

Typothorax
Temporal range: Late Triassic, 216–203 Ma
Typothorax coccinarum.jpg
Life restoration of two Typothorax coccinarum
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Aetosauria
Family: Stagonolepididae
Subfamily: Typothoracisinae
Genus: Typothorax
Cope, 1875
Species
  • T. coccinarum Cope, 1875 (type)
  • T. antiquum Lucas et al., 2002
Synonyms
  • Episcoposaurus Cope, 1887
  • Episcoposaurus horridus Cope, 1887

Typothorax is an extinct genus of typothoracisine aetosaur that lived in the Late Triassic. Its remains have been found in North America. Two species are known: T. coccinarum, the type species, and T. antiquum.

Typothorax was an aetosaur, a pseudosuchian distantly related to modern crocodilians. Unlike modern crocodilians, aetosaurs were herbivorous. Typothorax and other aetosaurs possess small, leaf-shaped teeth that were unsuited for a diet consisting of meat. Unlike some aetosaurs such as Desmatosuchus, Typothorax does not have large shoulder spikes. It does, however, have a pair of enlarged spikes on the neck projecting from the third row of scutes. It has lateral scutes that bear horns that are posteriorly hooked along its back, while its sides and underbelly are covered with ornamented scutes. Although fossils of aetosaurs are not as common as other Triassic archosaurs, with their armor plates being the most common, Typothorax has been represented by fewer skeletal elements than other aetosaurs. Typothorax was about 2.5 metres (8.2 ft) long and weighed 100 kilograms (220 lb).

The vertebral column of Typothorax is shortened, with individual vertebrae being reduced in length. However, the osteoderms that overly the vertebrae are not shortened. Instead, they are reduced in number so that each dorsal paramedian osteoderm (osteoderm that covers the back) overlies several dorsal vertebrae. In nearly all other crurotarsans, there is one row of osteoderms per vertebra. T. coccinarum has around 20 rows of presacral osteoderms and about 26 presacral vertebrae. If the cervical spikes of Typothorax are homologous to those of Desmatosuchus, it is likely that rows of osteoderms were removed from the front. This is because in Desmatosuchus the spikes are present in the fifth row, while in Typothorax they are present in the third.


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