A toothcomb (also tooth comb or dental comb) is a dental structure found in some mammals, comprising a group of front teeth arranged in a manner that facilitates grooming, similar to a hair comb. The toothcomb occurs in lemuriform primates (which includes lemurs and lorisoids), treeshrews, colugos, hyraxes, and some African antelopes. The structures evolved independently in different types of mammals through convergent evolution and vary both in dental composition and structure. In most mammals the comb is formed by a group of teeth with fine spaces between them. The toothcombs in most mammals include incisors only, while in lemuriform primates they include incisors and canine teeth that tilt forward at the front of the lower jaw, followed by a canine-shaped first premolar. The toothcombs of colugos and hyraxes take a different form with the individual incisors being serrated, providing multiple per tooth.
The toothcomb is usually used for grooming. While licking the fur clean, the animal will run the toothcomb through the fur to comb it. Fine grooves or striations are usually cut into the teeth during grooming by the hair and may be seen on the sides of the teeth when viewed through a scanning electron microscope. The toothcomb is kept clean by either the tongue or, in the case of lemuriforms, the sublingua, a specialized "under-tongue". The toothcomb can have other functions, such as food procurement and bark gouging. Within lemuriforms, fork-marked lemurs and indriids have more robust toothcombs to support these secondary functions. In some lemurs, such as the aye-aye, the toothcomb has been lost completely and replaced with other specialized dentition.