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Scale insects

Scale insect
Temporal range: Late NeocomianRecent
Scale insect.jpg
Adult female cottony cushion scale (Icerya purchasi) with young crawlers
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hemiptera
Suborder: Sternorrhyncha
Superfamily: Coccoidea
Handlirsch, 1903 
Families
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The scale insects are small insects of the order Hemiptera, suborder Sternorrhyncha. They comprise the superfamily Coccoidea, previously placed in the now obsolete group called "Homoptera". There are about 8,000 described species of scale insects.

Scale insects vary dramatically in appearance; from very small organisms (1–2 mm) that grow beneath wax covers (some shaped like oyster shells, others like mussel shells), to shiny pearl-like objects (about 5 mm), to creatures covered with mealy wax. Adult female scales are almost always immobile (aside from mealybugs) and permanently attached to the plant they have parasitized. They secrete a waxy coating for defense; this coating causes them to resemble reptilian scales or fish scales, hence their common name.

The group shows high degrees of sexual dimorphism; female scale insects, unusually for Hemiptera, retain the immature external morphology even when sexually mature, a condition known as neoteny. Adult males usually have wings (depending on their species) but never feed, and die within a day or two.

Species in which males do have wings generally possess only one pair of fully functional wings, and in particular, the forewings. This is unusual among insects; it most closely resembles the situation in the true flies, the Diptera. However, the Diptera and Hemiptera are not at all closely related and do not closely resemble each other in morphology; for example, the tail filaments of the Coccoidea do not resemble anything in the morphology of flies. The hind (metathoracic) wings of scale insects are reduced, commonly to the point that they generally are overlooked. In some species the hind wings have hamuli, hooklets, that couple the hind wings to the main wings, a condition usually associated with the Hymenoptera. The vestigial wings often are reduced to the point where they are referred to as halteres or pseudohalteres, but again, their resemblance to the halteres of flies is analogous, not homologous. It is not at present clear to what extent the pseudohalteres have any substantial control function to match the true halteres of the flies.


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