Nictitating

The nictitating membrane (from Latin , to blink) is a transparent or translucent third eyelid present in some animals that can be drawn across the eye for protection and to moisten it while maintaining vision. Some reptiles, birds, and sharks have full nictitating membranes; in many mammals, a small, vestigial portion of the membrane remains in the corner of the eye. Some mammals, such as camels, polar bears, seals and aardvarks, have full nictitating membranes. Often called a third eyelid or haw, it may be referred to in scientific terminology as the plica semilunaris, membrana nictitans or palpebra tertia.

Fully developed nictitating membranes are found in fish, amphibians, reptiles, birds and mammals, but are rare in primates. In humans, the plica semilunaris (also known as the semilunar fold) and its associated muscles are homologous to the nictitating membranes seen in some other mammals and other vertebrates. In most primate species, a plica semilunaris is present, although fully developed nictitating membranes can be found in lemurs and lorisoid primates.

Unlike the upper and lower eyelids, the nictitating membrane moves horizontally across the eyeball. It is normally translucent. In some diving animals, for example beavers and manatees, it moves across the eye to protect it while under water, and in these species it is transparent; in other diving animals, including sea lions, it is activated on land, to remove sand and other debris. This is its function in most animals. In birds of prey, it also serves to protect the parents' eyes from their chicks while they are feeding them, and when peregrine falcons go into their 200-mile-per-hour (320 km/h) dives, they will blink repeatedly with their nictitating membranes to clear debris and spread moisture across the eyes. In polar bears, it protects the eyes from snow blindness. In crocodiles, it protects their eyes from water but also hinders their focus under water. In sharks, it protects their eyes while the shark strikes at its prey. Woodpeckers tighten their nictitating membrane a millisecond prior to their beak impacting the trunk of a tree to prevent shaking-induced retinal injury.

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