Mesoraphidiidae
| Mesoraphidiidae Temporal range: Late Jurassic–late Cretaceous |
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| Amarantoraphidia ventolina holotype | |
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Raphidioptera |
| Suborder: | Raphidiomorpha |
| Family: |
†Mesoraphidiidae Martynov, 1925 |
| Subfamilies and Genera | |
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See text |
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| Synonyms | |
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See text
Mesoraphidiidae is an extinct family of snakeflies in the suborder Raphidiomorpha. The family lived from the Late Jurassic through the Late Cretaceous and is known from twenty-five genera. Mesoraphidiids have been found as both compression fossils and as inclusions in amber. The family was first proposed in 1925 by the Russian paleoentomologist Andrey Vasilyevich Martynov based on Upper Jurassic fossils recovered in Kazakhstan. The family was expanded in 2002 by the synonymizing of several other proposed snakefly families. The family was divided into three subfamilies and one tribe in a 2011 paper, further clarifying the relationships of the included genera.
Mesoraphidiidae are similar in overall appearance to modern snakefly species, having an elongated prothorax, giving a snake-like profile and from which the common name snakefly is derived. The family was likely tree-dwelling by nature, with larvae being active predators in trunks and branches. Female mesoraphidiids, where known and complete enough, have a long and densely annultated ovipositor which would have been used to lay eggs in crevices of bark. The family was restricted to the Northern Hemisphere, living in the warm temperate and paratropical forest belt that covered portions of Eurasia and North America.
As currently described, Mesoraphidiidae contains twenty-five genera, though several of the included genera were previously placed into separate families. As noted by Günter Bechly and Karin Wolf-Schwenninger in 2011, the defining characteristic shared by all members of the family is the placement of the forewings m1, m2 and m3 cells in a triangular pattern.
The genera Alloraphidia, Archeraphidia, and Pararaphidia were formerly placed into the family Alloraphidiidae. When reviewing the snakefly fossil record in 2002, paleoentomologist Michael S. Engel noted the lack of convincing distinctions between the two families, but tentatively retained the separation. In the same paper Engel moved the genera Huaxiaraphidia, Jilinoraphidia, and Sinoraphidia into Mesoraphidiidae and placed the monotypic families Huaxiaraphidiidae, Jilinoraphidiidae and Sinoraphidiidae as synonyms of Mesoraphidiidae. Engel did not give an explanation for the synonymizing of Huaxiaraphidiidae other than noting that the genus Huaxiaraphidia might be a synonym of Mesoraphidia and the type specimens should be reexamined. Similarly Engel did not give a specific explanation for the synonymy of Sinoraphidiidae beyond noting the poor type description of Sinoraphidia and a need to reexamine the type specimens. The creation of Jilinoraphidiidae was based on the apparent lack of crossveins in the forewings of the type specimen. However, Engel notes the lack of crossveins is due to the incomplete nature of the specimen, as there are several major longitudinal veins that show missing sections due to lack of preservation. The overall morphology and features of Jilinoraphidia are consistent with the other mesoraphidiids.
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