Litterfall, plant litter, leaf litter, tree litter, soil litter, or duff, is dead plant material, such as leaves, bark, needles, twigs, and cladodes; that have fallen to the ground. This detritus or dead organic material and its constituent nutrients are added to the top layer of soil, commonly known as the litter layer or O horizon ("O" for "organic"). Litter has occupied the attention of ecologists at length for the reasons that it is an instrumental factor in ecosystem dynamics, it is indicative of ecological productivity, and may be useful in predicting regional nutrient cycling and soil fertility.
Litterfall is characterized as fresh, undecomposed, and easily recognizable (by species and type) plant debris. This can be anything from leaves, cones, needles, twigs, bark, seeds/nuts, logs, or reproductive organs (e.g. the stamen of flowering plants). Items larger than 2 cm diameter are referred to as coarse litter, while anything smaller is referred to as fine litter or litter. The type of litterfall is most directly affected by ecosystem type. For example, leaf tissues account for about 70 percent of litterfall in forests, but woody litter tends to increase with forest age. In grasslands, there is very little aboveground perennial tissue so the annual litterfall is very low and quite nearly equal to the net primary production.
In soil science, soil litter is classified in three layers, which form on the surface of the O Horizon. These are the L, F, and H layers:
The litter layer is quite variable in its thickness, decomposition rate and nutrient content and is affected in part by seasonality, plant species, climate, soil fertility, elevation, and latitude. The most extreme variability of litterfall is seen as a function of seasonality; each individual species of plant has seasonal losses of certain parts of its body, which can be determined by the collection and classification of plant litterfall throughout the year, and in turn affects the thickness of the litter layer. In tropical environments, the largest amount of debris falls in the latter part of dry seasons and early during wet season. As a result of this variability due to seasons, the decomposition rate for any given area will also be variable.