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Halisaurinae

Halisaurinae
Temporal range: Late Cretaceous
Halisaurus BW.jpg
Halisaurus
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Sauropsida
Suborder: Scleroglossa
Infraorder: Anguimorpha
Family: Mosasauridae
Subfamily: Halisaurinae
Bardet et al., 2005
Genera

Halisaurinae is a subfamily of mosasaurs, a diverse group of Late Cretaceous marine squamates.

Bardet et al. (2005, p. 464) diagnosed the Halsaurinae as all mosasaurs more closely related to Halisaurus than Moanasaurus. Unambiguous character states were listed as follows: "premaxilla-maxilla sutural contact vertical anteriorly, oblique at midpoint and horizontal posteriorly; contact plane between the and the supratemporal oblique; preaxial ridge extending on two-thirds of the length of the radius; tibia and fibula long and slender with slightly expanded extremities. Ambiguous characters include "dorsal median ridge borne on the anterior two-thirds of the frontal; frontal with ventral boss; parietal foramen surrounded by a ventral boss; quadrate with large infrastapedial process; coalescent infra- and suprastapedial processes of quadrate; zygosphene-zygantrum complex absent; synapophyses of the cervical vertebrae extending ventrally to the ventral surface of the centrum; fused haemal spines."

Designation of this subfamily followed many decades of confusion surrounding the type genus, Halisaurus, especially H. sternbergii, a species from the Mooreville Chalk Formation of Alabama and the Niobrara Chalk of Kansas. Originally, the species had been referred to the mosasaurine Clidastes, then to Halisaurus (Russell, 1967; p. 369 ), which was also considered a member of the Mosasaurinae at that time. Later workers questioned the phylogenetic position and monophyly of Halisaurus, in part because of striking morphological differences between H. sternbergii and the other known species of the taxon. Finally, Bardet et al. (2004) determined that H. sternbergii was not conspecific with the other members of the genus and erected a new genus, Eonatator, as well a new subfamily, consisting of Eonatator and Halisaurus. Halisaurines (as members of this subfamily are collectively and informally known) were small to medium-sized mosasaurs averaging 4.5-6+ meters in length. Of all known mosasaurids, they were the least adapted to a marine lifestyle. Halisaurines are known from North America, Europe, South America, and Africa. The earliest known remains of halisaurines occur in rocks of Santonian age and the subfamily persists until the latest Maastrichtian. The etymology of this group derives from the genus Halisaurus (Greek halis = "sea" + Greek sauros = "lizard").


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