Greater short-toed lark | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Passeriformes |
Family: | Alaudidae |
Genus: | Calandrella |
Species: | C. brachydactyla |
Binomial name | |
Calandrella brachydactyla (Leisler, 1814) |
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Subspecies | |
see text |
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Synonyms | |
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see text
The greater short-toed lark (Calandrella brachydactyla) is a small passerine bird. The current scientific name is from Ancient Greek. The genus name, Calandrella, is a diminutive of kalandros, the calandra lark, and brachydactila is from brakhus, "short", and daktulos, "toe".
It breeds in southern Europe, north-west Africa, and across temperate Asia from Turkey and southern Russia to Mongolia. During migration they form large flocks that fly in synchrony and at other times they form loose flocks.
The greater short-toed lark was originally placed in the genus Alauda. The alternate name short-toed lark may also be used for three other species in the genus Calandrella. The Mongolian short-toed lark was formerly considered as a subspecies of the greater short-toed lark (as C. b. dukhunensis) until split in 2016 by the IOC. Formerly, some authorities also considered the red-capped lark to be either conspecific (as C. cinerea) with or a subspecies (as C. b. cinerea) of the greater short-toed lark.
Eight subspecies are recognized:
Several subspecies have been named but there is considerable geneflow and the species itself forms part of a larger complex. This is a small pale lark, smaller than the skylark. It is dark-streaked greyish-brown above, and white below, and has a strong pointed bill that is pinkish with a grey culmen. It has a pale supercilium, dark patches on each side of its neck and a dark tail. Some birds in the west of the range have a rufous crown. The sexes are similar. The greater short-toed lark is paler than the Mongolian short-toed lark which also has a shorter bill. In winter they fly in large and compact flocks that swing in synchrony. Care must be taken to distinguish this species from other similar Calandrella larks, such as the lesser short-toed lark.
The nominate form breeds in Europe (Iberia, France, Italy, the Balkans and Romania) and winters in Africa. Subspecies hungarica breeds in the eastern parts of Europe while rubiginosa breeds in north-western Africa. Subspecies hermonensis (sometimes including woltersi) breeds in Turkey, Syria and Egypt. Subspecies artemisiana (considered by some to be synonymous with longipennis) breeds in Asia Minor and winters in southern west Asia. Subspecies longipennis breeds in Ukraine, Mongolia and Manchuria and winters in South Asia mainly in the drier zone of north-western India.