Grammitid
| Grammitidoideae | |
|---|---|
 |
|
| Grammitis hookeri | |
| Scientific classification | |
| Kingdom: | Plantae |
| Division: | Pteridophyta |
| Class: | Polypodiopsida/Pteridopsida (disputed) |
| Order: | Polypodiales |
| (unranked): | Eupolypods I |
| Family: | Polypodiaceae |
| Subfamily: |
Grammitidoideae Parris & Sundue |
| Genera | |
|
See text |
|
See text
Grammitidoideae is a subfamily of the fern family Polypodiaceae, whose members are informally known as grammitids. It comprises a clade of about 750 species. They are distributed over higher elevations in both the Old and New World. This group was previously treated as a separate family, Grammitidaceae until molecular phylogenies showed it to be nested within the Polypodiaceae. It has since been treated as an unranked clade within subfamily Polypodioideae (renamed tribe Polypodieae in one classification), and, most recently, as a separate subfamily (reducing Polypodioideae to an evolutionary grade).
In 2011, Christenhusz et al. placed the grammitid ferns in the subfamily Polypodioideae, within the Polypodiaceae, as an informal group without rank. In 2014, Christenhusz and Chase expanded the circumscription of both family and subfamily, placing the Polypodioideae as previously delimited, including grammitids, in tribe Polypodieae. The PPG I classification of 2016 largely reverted to the 2011 circumscriptions of these groups, but placed the grammitids in a new subfamily.
The following phylogram shows a likely relationship between grammitid ferns and the Polypodiaceae subfamilies, based on Schuettpelz & Pryer, 2008.
"non-grammitid"
"grammitid" ferns
The grammitid ferns include the following genera. All except the most recent were formerly included in the family Grammitidaceae. Note that many of these genera have been found to be polyphyletic, so near-term changes to this list are likely.
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