In some egg-laying animals, the egg tooth is a small, sharp, cranial protuberance used by offspring to break or tear through the egg's surface during hatching. It is present in most reptiles, and similar structures exist in monotremes, Eleutherodactyl frogs, and spiders.
Some lizards develop a true tooth that is shed after use; other reptiles and birds generally develop an analogous epidermal horn that is reabsorbed or falls off.
A mother bird delivers her young encased in an eggshell; an external protective covering consisting of calcium carbonate (CaCO3). The shell protects the chick until it is ready to survive in the outside world. The chick breaks open the shell when it is strong enough and ready. Since the beak and the claws of the bird are not fully developed and cannot penetrate the eggshell, the "egg tooth" is the unusual structure that helps the bird break through the shell. It is only found in emerging chicks and lost soon after hatching, after it is used to penetrate the hard shell that once protected the embryo.
In birds, the process of breaking open the eggshell is commonly referred to as pipping.
Chicks have a pipping muscle on the back of their necks. It is this muscle which gives them the strength to force the egg tooth through the inner membrane of the eggshell.
When a chick becomes too large to absorb oxygen through the pores of its eggshell, it uses its egg tooth to peck a hole in the air sac located at the flat end of the egg. This sac provides a few hours' worth of air, during which the chick breaks through the eggshell to the outside. The egg tooth falls off several days after hatching.
Kiwis lack an egg tooth, instead using their legs and beak to break through a relatively thin eggshell. The superprecocial megapodes possess an egg tooth in their early embryonic development, but instead use their claws during hatching.
Baby snakes generally hatch from eggs with tough, leathery shells. A baby snake's egg tooth tears a hole directly through the shell, and falls off the first time the snake sheds its skin. Lizards have similarly leathery eggshells.