"Don't Ask, Don't Tell" is the 18th episode of the dramedy series Ugly Betty, which aired on March 22, 2007. The episode was co-written by Marco Pennette, Veronica Becker & Sarah Kucserka and directed by Tricia Brock. The episode's title references the United States' "Don't ask, don't tell" military policy.

In the wake of Claire Meade's arrest, the Suarezes find themselves dealing with the press coverage. The episode opens with a news reporter seeking to get a comment from Betty who, while trying to get away from the news cameras, runs directly into the wall of a bus shelter. Justin amuses himself by watching the footage repeatedly, which distinctly annoys Betty, who later expresses her guilt over Claire's arrest to Hilda.

At MODE, the now released Bradford and Alexis finally face off, with Bradford firing his daughter. Later on, Bradford goes to see Claire to see how she is doing. Claire insists that she has been better while he tells her that he is here for her. Daniel later comes in and Claire informs them that she wants them to be a family again, including Alexis. Bradford is extremely uncomfortable with the idea until Claire tells her husband that he can’t fire Alexis, since he does not own MODE. She reminds him that while he is the owner of Meade Publications, she owns the rights to MODE and therefore Alexis stays.

Meanwhile, Marc's mother — to whom he has never disclosed his homosexuality — visits the office; she brings along her cat, Lady Buttons of Camelot. Amanda learns that after three years of willingly pretending to be Marc's girlfriend during such visits, she has been "dumped" in absentia because Marc wanted to escape his mother's pressure to move on to marriage. Amanda gets angry and storms off, yelling to Marc that she will be the best 'pretend girlfriend' he has ever had.

Hours later, Betty tells Daniel that it may be a good thing that he and Alexis work side by side, but Daniel informs her that it was never easy working directly beside Alex, because he always had to come out on top. Daniel and Alexis later meet with Wilhelmina to inform her that they are both Editors-in-Chief at MODE and inform Wilhelmina that she is still the creative director. When he begins to claim what duties he will handle, Alexis interrupts and insists that she does not want anything to do with the business. After Daniel leaves the room, Wilhelmina rushes over to Alexis and tells her that she has to take what belongs to her and that she cannot leave MODE, then volunteers her office. Marc then panics when Wilhelmina drags him down to their new office and tells him it is only temporary—just long enough for Alexis and Daniel to push each other out.

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