Discosorida

The Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck.

The origin of the Discosorida is unknown, thought at one time to be directly from the Plectronocerida. Evolution within the order begins with the lower Middle Ordovician Reudemannoceratidae and from there diverges into three main lineages. Questionable discosorids have been reported as early as the Middle Tremadocian - near the start of the Ordovician, however the first bona fide examples date to the Middle Ordovician.

The diversification of the Discosorida, in terms of genera, peaked at the beginning in the Middle Ordovician (modern Darriwilian stage) followed by a decline in the Upper Ordovician (modern Sandbian and Katian stages) only to peak again in the Middle Silurian. Afterwards their diversity declined drastically and remained low until their end in the late Devonian. Some were endogastrically curved, with the lower, siphuncle side concave, others were exogastrically curved with the same side convex. In some, the aperture was a simple opening. In others, it became contracted into a pattern of slits. In earlier, Ordovician forms, the bullette became quite large and readily noticeable. In later forms, the bullette became reduced, in some to the point of being vestigial.