Cylons are a fictional race of robots in the original Battlestar Galactica TV series. They are the primary antagonists of the series and are at war with the Twelve Colonies of humanity. The Cylons also appeared in the short-lived 1980 spin-off series Galactica 1980.
An updated version of the original Cylon also appears in the re-imagined series: briefly in a flashback battle sequence in Battlestar Galactica: Razor, in hand-to-hand combat with a young William Adama after they had shot each other's fighters down; and several in the show finale "Daybreak", on board an old Cylon refuge known as "The Colony".
The Cylons were created by a reptilian race, also called Cylons. However, the reptilian race died out centuries earlier, leaving behind only their race of robots.
The Imperious Leader is the leader of the Cylon Alliance and most advanced Cylon model. According to the IL-Series Cylon Lucifer the Imperious Leader is an IL-Series Cylon, though it does not look like a IL-Series Cylon. The Imperious Leader has a third brain and a body shell resembling the reptilian Cylons. It is stated in the original novelization of the pilot episode that the Imperious Leader's third brain is specifically designed to emulate the human mind (solely for the purpose of anticipating human actions); although, since the novelization also specifically describes the Cylons as living, organic reptilian beings rather than robots, it's debatable whether the robot Imperious Leader has a third brain or not. One was killed at the Battle of Carillon (pilot episode) and his successor was possibly destroyed during the Battle of Gamoray (episode: "The Living Legend, Part 2"). In the original 1978-79 series, the voice of the Imperious Leader is identical to that of Count Iblis. (The voice of the former was provided by Patrick Macnee, who acted out the latter on-camera.) In the Galactica 1980 episode "Space Croppers," the voice of the Imperious Leader was provided by Dennis Haysbert. All of the Imperious Leaders look identical. The reptilian look of the Imperious Leader could have raised the question whether actually all living Cylons have died out.
Although they look much more civilian than centurions, IL-series Cylons act as commanders for the military and governors for civilians of the Alliance. They have two brains, and a mostly transparent head through which various lights can be seen pulsing. They also have a metallic, humanoid face with two eye scanners (compared to the single eye scanner of the Centurion models), and wear clothing (full-length glittering robes). Two IL-series Cylons have been featured onscreen, both of which have an effete human-sounding voice, unlike the flat mechanical tones of Centurions. They are: