Crepuscular

Crepuscular animals are those that are active primarily during twilight (i.e., the period immediately after dawn and that immediately before dusk). It is thus to be distinguished from diurnal and nocturnal behavior where an animal is active during the hours of daylight or the hours of darkness respectively. The term is not precise, however, as some crepuscular animals may also be active on a moonlit night or during an overcast day. The term matutinal is used for animals that are only active before dawn, and vespertine for those only active after sunset.

The time of day an animal is active depends on a number of factors. Predators need to link their activities to times of day at which their prey is available, and victims try to avoid the times when their principal predators are at large. The temperature at midday may be too high or at night too low, so for many varied reasons, crepuscular activity may best meet their requirements by compromise. Some creatures may adjust their activities depending on local competition.

The word crepuscular derives from the Latin crepusculum ("twilight"). Its sense accordingly differs from diurnal and nocturnal behavior, which respectively peak during hours of daylight and darkness. The distinction is not absolute however, because crepuscular animals may also be active on a bright moonlit night or on a dull day. Some animals casually described as nocturnal are in fact crepuscular.

Special classes of crepuscular behaviour include matutinal (or "matinal") and vespertine, denoting species active only in the dawn or only in the dusk, respectively. Those that are active during both morning and evening twilight are said to have a bimodal activity pattern.

The various patterns of activity are thought to be mainly antipredator adaptations, though some could equally well be predatory adaptations. Many predators forage most intensively at night, whereas others are active at midday and see best in full sun. Thus, the crepuscular habit may both reduce predation pressure, thereby increasing the crepuscular populations, and in consequence offer better foraging opportunities to predators that increasingly focus their attention on crepuscular prey until a new balance is struck. Such shifting states of balance are often found in ecology.

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