In zoology, copulation is animal sexual behavior in which a male introduces sperm into the female's body, especially directly into her reproductive tract. This is an aspect of mating. Many animals that live in water use external fertilization, whereas internal fertilization may have developed from a need to maintain gametes in a liquid medium in the Late Ordovician epoch. Internal fertilization with many vertebrates (such as reptiles, some fish, and most birds) occur via cloacal copulation (see also hemipenis), while mammals copulate vaginally, and many basal vertebrates reproduce sexually with external fertilization.
In non-primate mammals (rodents, canines, felines, bovines, and equines), the anatomy of the reproductive organs and some circuits of the nervous system are specifically organized for heterosexual copulation. On the other hand, human sexual intercourse stems from a change in biological factors that controls the copulation of mammals, and is mainly learned.
Spiders are often confused with insects, but they are not insects; instead, they are arachnids. Spiders have separate male and female sexes. Before mating and copulation, the male spider spins a small web and ejaculates on to it. He then stores the sperm in reservoirs on his large pedipalps, from which he transfers sperm to the female's genitals. The females can store sperm indefinitely.