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Caddisfly

Caddisflies
Temporal range: Triassic–Recent
Trichoptera caddisfly 1.jpg
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Hexapoda
Class: Insecta
Superorder: Amphiesmenoptera
Order: Trichoptera
Kirby, 1813

The caddisflies are an order, Trichoptera, of insects with approximately 7,000 described species. Also called sedge-flies or rail-flies, they are small moth-like insects having two pairs of hairy membranous wings. They are closely related to Lepidoptera (moths and butterflies) which have scales on their wings, and the two orders together form the superorder Amphiesmenoptera.

Caddisflies have aquatic larvae and are found in a wide variety of habitats such as streams, rivers, lakes, ponds, spring seeps and temporary waters (vernal pools). The larvae of many species use silk to make protective cases of gravel, sand, twigs or other debris. The name "Trichoptera" comes from Greek: (thrix, "hair") + (pteron, "wing").

Although caddisflies may be found in waterbodies of varying qualities, species-rich caddisfly assemblages are generally thought to indicate clean water. The caddisfly can be found in lakes, ponds, and marshes. Together with stoneflies and mayflies, caddisflies feature importantly in bioassessment surveys of streams and other water bodies. Caddisfly species can be found in all feeding guilds in stream habitats, with some species being predators, leaf shredders, algal grazers, and collectors of particles from the watercolumn and benthos.

Caddisfly larva with portable case of rock fragments

Caddisfly larva emerging from case made of plant material

Pupa of caddisfly

Close-up of spun case of caddisfly larva. BC, Canada

A "net" made by caddisfly larva. Texas, United States

Most caddisfly larvae are underwater architects and use silk, excreted from salivary glands near their mouths, for building. Caddisflies can be divided loosely into three behavioral groups based on their use of silk: net-making caddisflies and case-making caddisflies, both of which may enlarge their structures throughout their larval lifespan; and free-living caddisflies, which only make such structures prior to pupation. Net-making caddisflies usually live in running water, and their nets, often made amongst aquatic vegetation, serve both as a means to collect algae, detritus, and animal food and as retreats. Case-making caddisflies may build cases exclusively of silk, but more commonly the silk holds together substrate materials such as small fragments of rock, sand, small pieces of twig or aquatic plants.


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