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Arthropleuridea

Arthropleuridea
Temporal range: Late Silurian - Pennsylvanian 423–299 Ma
Arthropleura 1914.jpg
Arthropleura fossil illustration
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Myriapoda
Class: Diplopoda
Subclass: Arthropleuridea
Orders

Arthropleuridea is an extinct subclass of myriapod arthropods that flourished during the Carboniferous period, having first arose during the Silurian, and perishing due to climate change just before the Early Permian. Members are characterized by possessing diplosegement (fused "double segments", as in modern-day millipedes) paranotal tergal lobes separated from the body axis by a suture, and by sclerotized plates buttressing the leg insertions. Despite their unique features, recent phylogenetic research suggests Arthropleuridea be included among millipedes in the class Diplopoda. The subclass contains three recognized orders, each with a single genus.

Arthropleurids lived in the moist coal swamps that were common at the time and may have burrowed in the undergrowth. They were either herbivores or detritivores. Besides their size, their most distinguishing features were their legs with eight segments (as many as 30 pairs) and extremely tough exoskeletons. There is no evidence of spiracles, so the animals must have used lungs or gills for respiration.

Most arthropleurideans are thought to have been terrestrial, although, without any known respiratory structure, terrestriality is assumed only by analogy to modern arthropods. Early forms, however, including Eoarthropleura (order Eoarthropleurida), appear to have been aquatic. For this reason, some question Arthropleuridea's inclusion among millipedes because no modern aquatic myriapods are known.


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Wikipedia

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